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Well physiologically the biceps have almost nothing to do with a punch, and those are his biggest assets. His pec(toral)s, the main muscle used in a punch, are not the strongest looking either( I’m not saying I am better, I’m a twig ). Furthermore, he is likely mostly slow-twitch muscle, which is not great in a fight but great for chasing down prey. I guessed this as slow twitch muscle is larger than its fast twitch counterpart, and his biceps are massive. (sorry for the anatomy lesson).